lec 1 Flashcards

(53 cards)

1
Q

What is the word evolution derived from

A

latin word Evolutio

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2
Q

what does the latin word “evolutio” mean

A

unravelling or unfolding

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3
Q

What is the quote by Dobzhansky in 1973

A

Nothing in biology makes sense except in light of evolution

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4
Q

What is evolution

A

descent with modification;

the earth was made 4.5 bya and the first life forms came 3.7 bya

Our closest ancestor was chimpanzee- but we didnt eolve from chimps we just both have the same common ancestor that lives 5.5 million years ago and there was descent with modification

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5
Q

What isnt evolution

A

it does NOT include development change in the life of an organism
and IT DOES NOT PROCEED ALONG SOME PREDICATABLE COURSE

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6
Q

TOPIC 1

A

History of the development of our modern understanding of evolution

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7
Q

WHO proposed that populations of organisms change over time

A

1) Anaximander (ancient greece, 6th century BCE)
-thought species were formed from water and humans and other animals came from fish

2) Empedocles (5th century BCE)
-thought head, limbs and other organs were joined at random and only some combos were fit for survival

3)Plato (4th century BCE)
-each natural type of object in the observed world is an imperfect manifestation of the ideal (ie there is an ideal structure for tings but what we are is the imperfect)

4) Aristotle (student of Plato and elaborated on his ideas)
-envisioned static world in which a deity created “Fixed essences” which were templates for all species
-believed species were unchanging

Scale of Nature (Scala Naturae)
-graduation from inanimate to barely animate through plants and invertebrates up to higher forms
-moves from least perfect to most perfect
-Christianity expanded on this : GOd’s creation follows a plan - the lader of nature or Great Chain of Being
therefore: aristotle beleives SPECIES ARE PERMANENT AN UNCHANGING SINCE CHANGE WOULD IMPLY IMPERFECTION IN THE ORIGINAL CREATION

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8
Q

what is aristotles view of species

A

spcies are permanet and unchanging cuz change would imply imperfection in the orgiinal creation

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9
Q

Where did species orginate from?

A

THEORY OF SPECIAL CREATION: Archbishop James Ussher
1) Created by a deity
2)Species were created independetly of one another
3)Species remain UNCHANGED since their creation
4) variability within species is minor/ limited (fixe static creatures, any variation is unfortunate)
5) Predicted that creation event occured 26 october 4004 BC at ( (6000 years ago only)

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10
Q

When does the theory of special creation think that creation occurred

A

october 26 4004 bc (only 6000 years ago)

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11
Q

who predicted based o the bible that the earth was made oct 26

A

archibishop james ussher

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12
Q

Special Creation vs Descent with modification

A

Special Creation:
1) Species are unchanged
2) Lineages do not split/diverge
3) All species are created independelty
4) All species are created seperately
5) Earth is young

Descent with modificaton
1) Species are changing. (microevolution)
2) Lineages diverge and split (speciation)
3) New life forms are derived from old ones (macroevolution)
4) all life forms are connected (Common ancestory)
5) Earth an life are old

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13
Q

Carolus Linnaeus (1707-1778)

A

-Swedish naturalist
-Father of Modern Taxonomy
-Classified plants based on morphology into KINGDOM, CLASSES, ORDER , GENUS , SPECIES (missing kingom, domain, phylum)
-introduced a system of binomial nomeclature for species identification (systema naturae 1735)
-individual species names consist of genus and species name
-remains standard today
-ie Homo sapiens
-established framework for modern classification (the binomial system) in Systema naturae in the hope of discovering God’s pattern of creation (if we can understand species, we might understand god’s creation)

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14
Q

Where did species orginiate

A

Theory of Evolution: Descent with Modification

Erasmus Darwin:
-Laws of Organic Life (1794-1796) , Integrative approach
-he was familiar with animals in captivity and artifical selection of niamls in captavitiy,
-he determined: Final course of contest among males be that the strongest and most active animal should propogate the species which should thus be improved (the ones that have access to females were the strongest and most active)

Jean Baptiste Lamarck:
-Wrote Pbilosophie Zoologique (1809)
-Lamarckasm

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15
Q

What is lamarckism

A
  • All species are derived by GRADUAL change from other species
    Mechanism: Inheritance of acquirred characteristics
    -mechanism required that bodily changes have a way to be registered int he germ cell
    -“organs become improved if used alot, reduced to vestiges if not”
    -ie giraffe and necks, the prodgency will be more stretcjed and stretched
    ie 2: Humpback whales: have hindlimb vestiges
    ie3: Black-necked stilts (long legs, strethcing to stay above the waterline”)

is this right:
-NO because under this way of thinking, a bodybuilder would have kids with bigger muscles ie arnold schwartnegger

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16
Q

Theory of Geological History of Earth

A

2 theories about the hstory of the earth and how it is young and h0ow it is: CATASTROPHISM VS UNIFORMITARIANISM

Catastrophism:
-biblical interpretation of the hisotry of earth
-earth was approc 6000 years
-had been affected by a series of catastrophical events such as the biblical flood

Uniformitarianism:
-more modern view of eaths history
-earth is very old
-landforms are present as a result of geolocial processes that are stil occurring (ie volcanoes, erosions,)
-ideas originated from the work of James Hutton 1700s
-father of modern geology

Charles Lyell:
-Published “Principles of geology (1830-1833) IMPORTNAT
-challenged the belief regarding earths history and age
-argued earth is hundreds of milions of years old
-earths geology was created by natural forces INCLUDING earthquakes, volcanos, winds, race
-charles was looking at the impacts of eorsion and how they cahanged earth (looked at this in england)

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17
Q

Charles Darwin

A

-born 1809 after lyell
-dropped out of medical schools after 2 years at 18
-stidoed fpr tje clergy at Cambrodge where he developed interest in biology
-graduated in 1831 (began working with John Hindslow at C uni)
-accepted unpaid position as nautralist aproad HMS beagle (British Navy Mapping Expedition)
-took lyells “principles of geologies” book on the expedition

Voyage of Beagel: 1831-36

Observations from South America:
1) Fossils of species that were no longer extant in the region
ie THoatherium (horse like speices, no horses in south america) THEREFORE species can and will go extinct (so if a deity made, they dont all exist)

2) Fossils of animals in a region were different to modern day equivalents (ie armadillos) ; ancestor was glyphodont
-so species have ancestors, and CHANGE through time

3) Travelling from North to Sourth, one species was replaced by a similar, but slightly different species
ie RHEAS (rhea americana different from rjea darwini)

Galapagos islands:

TORTOISES
-isolated from other islands
-tortoises were present on every island, but each island had toroties sligtly different from the other ones
-14 species of tortoises that lived in the galapogos islands
-some are extinct now because sailors used to eat totoises , and would bring them on ships and they could kill ir later and have fresh meat
-DODO bird became extinct quick from maritious islands because they were “dumb” and didnt fly cuz no predatirs, but when humans showed up they ate them and dodo birds didnt exist anymore

DARWIN’s FINCHES:
-all finches, all different on different islands
-each have diff body size, beak size, beak shape,
-suited for aquirring different foods and insects and fish
-for fish: large gil, for insects: small pointed gill

-goerge couvier, documented 20 species tht went extinct
-darwin believed what he was saying
-everyone else thihgt that they hadnt found them yet but darwdin didnt agree
-ie Irish elk

therefore:forrisl records told us

a) epcies go exticnct
2)L aw of sucession
3) transitinal forms
-if we have brids that look like crocodile, we must beable to find transitional forms in fossil regords
-ie archaeopetry lithographica (first ancestral bird, has teeth, 3 toes, claws, -birds and dinosaurs
4) environmental chsnge: found fosssils that were clearly of terrestial speic,es they were marine animals, HOw could that have been?
-theere are sepcies that come from land and water

BACK AT HOME IN ENGLAND:
a)
Vestigial organs: -darwin figured this out in england
-“Functionless or rudimentary versio of a body part that has an important function in a closely allied species

ie Mexican Tetra: developed with tiny vestigial eyes under skin
-eyes didnt develop from ancestors anymore because ancestors maybe lived in darker areas so they ddint use their eyes anymore so now in enctant species there is no eye present
ie 2: Brown Kiwi : have tiny, useless wings, burried under feathers and cant fly
WHY: ancestors were flying birds that got to new zealand and had no predators so didnt need to fly anymore, smaller wings saved energy

ie 3: Rubber Boa: have small spurs near tail that is like hindlegs , pelvis
-why: developed from four legged ancestors that began burrowing and slithering so legs hd no benefit anymore

b) Artificial Selection:
-humans have modified many species to suit their needs thorugh careful breeding programs
ie SHETLAND PONY, ARABIAN HOURSE, plants
so humans have “Caused” the evolution of several species from a single ancestral species
-many domestic fowl came forom red jungle fowl…
but what was the mehcniasm of inheritance?

c) in 1838, read the principle of population by THOMAS ROBERT MALTHUS (1766)
-discussed how reproducive capaciry of rhe human populatio EXCEEDS fod supply available to noruish the growing population, therefore humans compete among themselves
-this struggle would also exist in nature and constitue a natural selection process that occurs without plan or conscious
-darwin prepared an essay of ideas in 1844 and showed LYELL who urged him to make book but darwin only published ideas much later

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18
Q

La of succession

A

correspondence between fossils on area, and descended frm here there are new species that are slightly different’-same place, over time we see changes in seocies

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19
Q

SO what are examples of animals with vestigial structures

A

1) Mexican Tetra
2) Brown Kiwi
3) Rubber Boa

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20
Q

Homology

A

-study of likeness
“the same organ in diferent animals uner every variety of form and function “ R. Owen

IE FORELIMBS:
-all have radius, humerus, ulna, carpus (wrist), phalanges but all are organized in different form to suppoort the animal
horses: didnt have phalanges bevause use was for running
Dolphin: longer phalanges to help with swimming and to become the “fins”
Bat: each phalange became parts to support the wing shape
mole: used for digging, so the forlimb became stubby and thicker

Traits are homologus if they are derived, evolutionarily and developmentally from the same source structure

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21
Q

Who is the principles of population by and what did it say

A

THomas RObert Malthus

darwin read in 1838

said: the reproductive ability of the human population is more than the food supply to nourrish the expanding population and therefore, humans compete with eachtoher
-this struggle would also exist in nature and constitue a natural selection process

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22
Q

Alfred Russel Wallace

A

1823-1913
-working class family
-father died young
-worked with brother as surveyor
-met Henry Walter Bates who introduced him to BOTANY
-both him and henry walter bates travelled to South America and collected for 4 years
-lost all their collections in fire uring return vogage in 1852
-also travelled to MAlay Archipelago (12 months)
-read essay by malthus and also came to nautral selecton idea during malaria fever
-Wrote “ on the tendencies of varities to depart indefinitely from the orgfinal type” and sent for review from darwin

23
Q

Linnean Society of London

A

July 1 1858
-lyell ad hooker arranged to share their ideas of darwin and wallace
called
“on the Origin of Spcies by Mean so fNatural Selcetion, or the preservation of favoured races in the struggle for life” Nove 24 1859,

What is the Linnean Society of London: founded 1788
-forusm fro disscusions about genetics, natural history, systematics, biology, and history of plant and animal taxonomy
-worlds oldest extant biological society
Named after Carl Linneasu

24
Q

Darwins Concept of Natural Selection

A

Observation 1: Potential exponential increase of population: 1 single elephant can make 19 million in 750 years if all optimal conditions and all progeny live

Observation 2: Steady stae, ie stability of populationsL this is the reality of how life goes, even though many are born, many die
-deduction 1: Struggle for existence among individuals: ie fight for resources

Obseration 3: Heritable variation among organisms: individuals are diferent
deduction 2L Differential survival: natural selection

25
What could Darwin Not explain
1) how a given traist is transmitted from parent to offspring 2) Where and how new traits suddently emerged GREGOR MEDEL: -Conducted experiments (1856-1863 !!!!!) in plant hybridization using common peas and was credited for laws of heredity (Basics for all meodern genetics) -signficiance of his work not realized until 1900
26
Gregor mendel pea experiment
Pisum sativum -used pod shape, colour, and seed shape colour and flower colour to figure this out Discovered 1) Traits are passed from parent to offspring through gametes in specific discrete units or factors -these factors can be assorted differently in different indivduals -factors are GENES now 2) Individuals have 2 copies of each gene -one each parent -alternate forms of the genes are called ALLELLES What are the principles `1) Principle of Segregaton -each gamete can only carry one allele from a parent at a given gene 2) Independent assortment -segregation of members of one pair of alleles occurs INDEPENDETNLY of the segregation of the members of another pair of allele in gamete formation
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lec 2
Evidence and patterns of evolution
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Orgin of species by means of natural selection was presented at lineans in
1858 published in 1859
29
when did darwin have his first phylogenetic tree
1837
30
when was charles darwin on the beagle
1830s
31
up until what year was it illegal to teach evolution in class in US
1967; when the book was written 1859 in other countries alike japan and europe had more acceptance for evolution iceland had the most acceptance for evoution whereas USA did not only 40% in uSA thought it was true that humans developed from earlier species of animals After 2007, there was an increase in the acceptance if evolution after release of paper in2006
32
What is a species
Biological first definition: "one species never springs from the seed of another"
33
Evidence for Descent with modification
a) Species do change (microevolution) Selective Breeding (ie artificial selection) -you put mice on wheel: some run fast, some run slow, some run for long time -measured how long mice ran voluntarily, and number of revolution per day -created 8 populations of mice: 4 fastest (highest revolutions per day), and 4 at random -mated 4 pairs of fast and fast -did for 24 generations then measured how much the offspring run : average 14000 revlutions for the fast offspring, for the random: average was 5k -SUGGESTS: by seelcting the individuals that ran alot, they were able to cause microevolution in the population in over 24 generations -the individuals didnt EVELOVE, the population evolved Vestigial Structures: a useles or rudimentary structure or version of a body art that has an important funcition in other closesly allied species, cannot be explaine by theory of special creation but makes sense for theory of descent (mexican tetra (eyes), brown kiwi (Wings), boa (legs), humans (tail bone, arrector pilli muscle), Threespine Stickleback (marine vs freshwater armor, test) b) lineages do diverge (SPECIATION) Speciation is: the concept that describes the situation where new species arise from pre-existing species -starts from a single population where there is variation among individuas What is a species: Biological Species Concept: Species are populations, or groups of populations within and among which individuals actually or potentially interbreed, and outside of which they do not interbreed (ernst mayr) Example 1: Threespine Stickleback Example again: Single population with 3 variations: blue, purple, green, all can interbreed with eachother because same species -over time, they divide more inot subpopulations (can still interbreed but more distinguishable) -over more time: we get distinct populations with limited interbreeding (still can but much less often) -finally, become reproductively isolated populations = different species So basically u can have a population with variation and it will al still be the same species, but over time, if they become more isolated, then they will become distinctly different species Example 2: Ring Species (siberian greenish warbler) -Ring species: series of populations of a species that geographically arrange around a barrier like a mountain or lack of forest -adjacent populations may be able to interbreed, but geogrraphically seperated ones wont be able to For Siberian Greenish Warbler: -differences in genetics and song (mating call) is strong for populations that are more geographically isolated (they do not recognize eachtoerhs mating call) -shows how they are slowly reproductively isolated c) new life forms are derived from older life forms (MACROEVOLUTION) - Goerge Cuvier: Published list of species only known from fossils (1801) -Clift noted that extinct mammals from australia were like extant marsupials (Slot, irish elk) -darwin noted the same things with sloths and armadillos in south america -SHOWE THAT fossils and living org in the same geographical area resembled eachother and are distinct from organisms elsewhere (LAw of Succession) EXAMPLR : Transitional Forms: Pacific Leaping Blennies: Transitional meaning u can see the transition states in evolution Aquatic ones: cant leave water Amphibious ones: double life -there was a point in time where a common ancestor that was aquaitc transitions to amphibious and from there there was a transition to Teresstial: can persist in water but for short term, will jump out, preferes moist environment -can survive on land , and also respire across gills, o2 crosses and enters blood stream Example 2: What is a bird : Archaeopteryx and phylogeny: -Bird: flight and feathers -but share many traits with crocodiles and dinosaurces because there is a common ancestor of all of these -vestigial features are seen on sinosauropteryx (short feathers that are no flight feathers but used for insulation and thermal regulation) -similicaudipteryc : long feathers, more similar to birds but not for flight From Dinos to birds: sinosauropteryx and similicaudipteryx d) All life forms are related (common ancestory) -All organisms have a common ancestor -Richard Dawkins "the Ancestor's tale". -Humans are connected all the way back to Protocells/unicellular life -protocells -> multicellular life - > animals - > fish and tetrapods -> malla s -> great apes -> and homo sapiens (NOT THE END THUOGH< evolution is still happening) -Homologoy is crucial line of evidence : traits are homolougs if they are derived evolutionarily, and a developmentarily from the same source structure. (ie 4 limbs) A) Structural Homology: -each structures adapteed to specific environemtns (Whale, horse, humans,moles, bat) -the genes that control the evelopmentof these is the same but they are different in function and stucture Darwin and Orchids Analogus: similar function but derived volutionarily and evelopmentally from a DIFFERENT source structure ie whales and sharks : whales have strong tail from terrestiral form, sharks from aquatic Molecular Homology: -Flaw shared between humans and chimpanzees and bonobos but not gorillas , orangutans or others -chimps , humans and bonobos all have an error in their gene , showing that they inherited this gene error from a commmon ancestor -both have cahrcot-marie-torth disease and hereditary neuropathy (pressive palsies) -Finally : the fact that all species have a universitality of genitic code and triplet nucleotides that code for AA (codons universal) e) Earth is old Catastrophism (biblical) / univormitarianism -alot of time was needed to build mountains and create the geoloical stata observed in nature - based on evidence, 6000 year old earth (arch bishop usher) and catastrophism is not consistent with evidence -calendar divded into Eons, Eras, Periods, Epochs
34
DO individuals evolve
NO populations do
35
Whar is the definition of a population
group of same species that live in a specific area at a specific time, so that the individuals have the potential to mate with eachtoher OR -most cases: there are multiple populations of a different species (several independent moose, or galapagos turtles (darwin))
36
Another efinition of evolution
change in alellele frequencies in alele frequency in a population over time -for the mice, we measure the traits and it changed in the popylation over time, so it is the population evolving
37
What is the common ancestory of all tetrapods (four feet)
Tetrapoda (four feet)
37
Vestigial structures are homologous traits?
Usually becaue homologous if they are derived, evolutionarily and evelopmentally from the same source
38
Vestigial Structure of HUmans
Tailbone -dont have tails but have this Errector Pilli Muscles -next to hair follicles -think about cats, when scared or threatened, hairs stand up, it is a behavioural response to threat -for these animals, the ahir is also insulation, not for us Thermal Regulation: we and chimps are homeotherms (we maintain the same body temp regardles of external temperature), if my body temp drops, it means we will be sick -homeotherms have adaptations to maintain temps then -our hairs still stand up but it doesnt provie thermal protection but we still have this thing but for other mammals the hair standing up provides warmth and insulation
39
Threepine stickleback Vestigial structures
ancestral form is marine: these are fully armored with pelvic fins modified into spines FreshwaterL Ligher armor, vestigial pelvic structure both are from one genotype, each is a different allele -each trait controlled by a single gene, with two alleles (one for light armour, one for heavy) Test: -all fish in Lobery lake Alaska were poisoned in 1982 for restocking -lake was colonized by threespine sticklebacks from marine inlet (1988) -from 1990-2001, freuqencu of individuals with ifferent traits forms recorded Results: Lightly plated increased in population, and armour decreased. (POPULATION CHANGED DRAMATICALY) -individuals didnt evolve, but POPULATION CHARACTERISTICS DID Why did the fuly armoured ecreased: less predators, so decrease in this phenotype -shows us how the less armoured are modified descendantsof marine populations NOTE: Need does not drive evolution or change : For change to happen, gene variants need rto have already existed, an only the frequency of this changes (lightly plated already existed ie)
40
Microevolution vs macro
micro : over generations' macro: more major changes
41
What are the animals and their examples for
Vestigial Structures 1) Mexican tatra (Eyes) 2) Brown Kiwi (wings) 3) BOa (hindlimb) 4) Humans (tailbone, arrector pilli muscle 5) Threepine stickleback (armour and pelvic fin) Microevolution: Vestigial features and threepine stickleback Speciation: Threepine stickleback and 3 coloured variation thing b) Ring species (Siberian greenish warbler) MAcroevolution: ex 1: Pacific leaping blennies : they are the transitional state between fully awuatic and fully terrestrial. -show evolution because live double life ex : 2 What is a bird : Archaeopteryx and phylogeny: -Bird: flight and feathers -but share many traits with crocodiles and dinosaurces because there is a common ancestor of all of these -vestigial features are seen on sinosauropteryx (short feathers that are no flight feathers but used for insulation and thermal regulation) -similicaudipteryc : long feathers, more similar to birds but not for flight From Dinos to birds: sinosauropteryx and similicaudipteryx
42
Homology vs Analogy
Analogus: similar function but derived volutionarily and evelopmentally from a DIFFERENT source structure ie whales and sharks : whales have strong tail from terrestiral form, sharks from aquatic Molecular Homology: -Flaw shared between humans and chimpanzees and bonobos but not gorillas , orangutans or others -chimps , humans and bonobos all have an error in their gene , showing that they inherited this gene error from a commmon ancestor -both have cahrcot-marie-torth disease and hereditary neuropathy (pressive palsies) -Finally : the fact that all species have a universitality of genitic code and triplet nucleotides that code for AA (codons universal)
43
Earths history in a single year
-muticellular organisms appear in the ocean late June -Land plants appears late november -the swamps that formed the world's coal deposits during the pennsylvanian thrived for 4 days in dec -dinos: mid december, but extinct boxing day , rocky mountains formed -human-like animals appear in evening of dec 31 -most recent galciers receded from waterloo area 75 seconds before midnight on 31st -the Roman empire ruled western world for 5 seconds, from 11;59:45 ro 59:50 on 31st -Columbus arrived 3 seconds before midnight
44
Lecture 3
Natural Selection
45
What are the KEY WORDS/ COMPONDENTS in the definition of Evolution
1) Change/MOdification 2) Genetics/Allele frequencies 3) Population 4) Time
46
What are the mechanisms that cause change in alelle frequence over time (mech of evolution)
1. Natural Selection 2. Mutatino 3> Gene flow 4. Genetic Drift 5. Non random mating
47
Theory of Evolution by Means of Natural Selection (Darwinian evolution and wallace)
All must be true for nautral selection: 1) Individuals of a population differ from one another (if everyone is a genetic clone, natural selection cant happen) 2) THe differences are , at least in part, passed from parent to offspring (phenotypical differences) 3) Some individuals are more successful at surviing and reproducing than others 4) The more sucessful individuals are not just lucky, hey suceed bebcause of variant traits they have inherited an will pass to offspring In another way: a) organisms produce more offspring that can be supported _as population size increases, the amount of food goes down, then survival and reprouction become compromised - as more food available, it increases population size, so less food, so less growth b) There is constant struggle for existance c) Invidiaus in population vary in phenotypes d) Some of this phenotypic variation is heritable (genotypes) e) Those individuald that have phenotypes that are most adapted to current conditions are most lkely to surive and reproduce f) if rthese are heritable, it will be passed on to offspring Natural selecrion acts deirectly on phenotypes of individuals, but only if there is a change in allele frequencies in a populat occurred If all of these postualiteds hold true, then population will change gradually over time Evolution in population results from natural selection on individuals; requires change in frequency distribution of a trait in population (statistical thinking)
48
What is population genetics
study of allele and genotype frequencies and the variation in these frequences acrosss space and through time and evolutionary factors that explain this variation
49
What is darwinian evolution
if all the autral selection postualates hold true, then populations will gradually change over time evolution o a populatinn results from natural selection on individuals
50
Loci vs Gene vs Alleles
51
Natural selection acts on phenotypes of individuals but cumulative effects of natural seelction of all indiv in population, results in changes of population from gen 1 to 2 to 3 etc
these generational changes are usually small and it might take hundreds of generations to detect meaningful difference
52
What is fitness
ability of an organism to survive and reproduce in its environment -relative term, relatice to the fitness of other members of the population -lifetime reproductive stress