What is a receptive field?
Area of retina where light changes the activity of a neuron (bipolar or ganglion cell).
What forms the centre of a bipolar cell’s receptive field?
Direct input from photoreceptors.
What forms the surround of a bipolar cell’s receptive field?
Indirect input via horizontal cells.
Why does the surround have the opposite effect to the centre?
Horizontal cell inhibition → enhances contrast detection.
In darkness, what do photoreceptors release?
More glutamate.
How does an ON-centre bipolar cell respond to light in the centre?
Photoreceptor hyperpolarises → glutamate ↓ → ON bipolar depolarises → ganglion firing ↑.
How does an ON-centre bipolar cell respond to light in the surround?
Photoreceptor hyperpolarises → horizontal cell inhibition removed → ON bipolar hyperpolarises → ganglion firing ↓.
How does an OFF-centre bipolar cell respond to light in the centre?
Photoreceptor hyperpolarises → glutamate ↓ → OFF bipolar hyperpolarises → ganglion firing ↓.
How does an OFF-centre bipolar cell respond to light in the surround?
Photoreceptor hyperpolarises → horizontal cell effect → OFF bipolar depolarises → ganglion firing ↑.
What do centre–surround receptive fields specialize in detecting?
Edges and contrast, not absolute brightness.
Why do we perceive objects mainly through contrast?
Visual system compares object to background, not raw light intensity.
What layers of LGN receive rod (magno) input?
Layers 1–2 (magnocellular).
What layers of LGN receive cone (parvo) input?
Layers 3–6 (parvocellular).
Do the eyes mix before cortex?
No — left/right eye input remains separate in the LGN.
Where does LGN project in V1?
Layer 4C of primary visual cortex.
Which V1 sublayer receives magnocellular input?
4Cα.
Which V1 sublayer receives parvocellular input?
4Cβ.
What are ocular dominance columns?
Alternating columns responding to the left or right eye.
What are orientation columns?
Columns of neurons tuned to a specific orientation of a bar or edge.
What do simple cells in V1 detect?
Orientation + position, responding to bars/edges in exact locations.
How do complex cells differ from simple cells?
Same orientation tuning but position-invariant; respond best to moving edges.
What does area MT (V5) process?
Object motion.
What does area MST process?
Linear, radial, and circular optic flow (self-motion).
What does V4 process?
Colour and shape (orientation + colour-selective).