Mollusca
a lot of them are seafood
- oysters, snails, squid, mussels
also used in architecture and art
Adaptive Radiation
adaptive radiation = rapid evolutionary changes within one lineage that produces a lot of descendent species with a wide range of adaptive forms
evolutionary radiation = pattern of evolution where multiple divergences from a common ancestor in a short period of time happened
Radiation History
Molluscs went under radiation around 600 million years ago
- 8 descendant lineages (classes) survived
4 lineages:
- Polyplacophora (chitons)
- Gastropoda (snails and slugs)
- Bivalvia (clams, oysters, mussels)
- Cephalopoda (squids, octopuses)
***mollusca and annelida are most closely related
tree:
- platyhelmines, then mollusca and annelida
Mollusc Traits
Bilateral symmetry, triploblast, coelomate, unsegmented protosotomes (spiralia)
Plasticity
the molluscan body plan has high plasticity
- the body plan can be molded like a plastic so it can fit different ecological niches
***mulluscs are very successful because they have extreme plasticity and can adapt
Molluscs Model
They aren’t segmented
- but, they are made of some basic unit that can be changed in the same way segments are
Basic Parts of Mollusc
basic parts are the ventral foot, dorsal visceral mass, mantle, and shell
Class of Mollusc: Bivalvia
two shells that are dorsally hinged
- suspension feeder
Class of Mollusc: Polyplacaphora
broad flat foot and shell is divided into 8 plates
Class of Mollusc: Gastropoda
most have coiled shell
- well defined head and foot for crawling
- largest class of molluscs
Class of Mollusc: Cephalopoda
free swimming
- have very developed nervous system
- predator
- their foot is modified into tentacles
- most don’t have a shell
Coelom and Hemocoel
coelom is reduced
- it exists only in fluid filled space surrounding a heart
hemocoel = principle body cavity
- made of blood-filled sinuses (spaces) surrounding the internal organs
- homocoel is the functional hydrostatic skeleton
Ventral Foot
the ventral side (underside) is made of a broad, flat, muscular foot
- the muscular foot is involved in locomotion (movement)
Head with Radula in the mouth
the radula is a ribbon of teeth
- the teeth are made of chitin and placed in rows
- they move back and forth by set of muscles
- chitin = polysacharide
radula is feeding structure located within the mouth cavity:
- radula is pulled over food particles (think of algae growing on rocks)
- this process breaks off piece of food and bring them into the mouth
Dorsal Visceral Mass
the internal organs are concentrated in a region on top of the of the foot called the dorsal visceral mass
Mantle
sheet of skin that is located between the visceral mass and the shell
*the mantle secretes the shell
Mantle Cavity
the space between the outside wall of the visceral mass and the mantle = mantle cavity
- this space is open to the environment
the digestive system empties waste into the mantle cavity via the anus
- the kidney and gonads also empty their products into the mantle cavity
Gills
the gills are covered with cillia that create a current that draws water into the mantle cavity and over the gills
Shell
made of calcium carbonate
- deposited in a framework of protein
Nervous System
has a ganglion that circles the esophagus and nerve cords extend to the organs and foot
Movement
the hemocoel is the hydrostatic skeleton
- the muscles contract in waves that work against the hemocoel, which produces movement
Open Circulatory System
the blood is pumped by a heart into vessels (blood leaves the vessels)
- the vessels come to an end and the blood flows into the spaces or cavities around the cells
hemocoel = cavity around the cells where the blood flows
- gases and nutrients exchange between the cells and the blood in the hemocoel
*blood flow is slow in open circulatory system because it empties into a big space and the pressure can’t be maintained
- less efficient than closed circulatory system
Getting Nutrients
complete digestive tract:
- made of mouth, esophogus, stomach, intestine, anus
***in the intestine, digested nutrients are absorbed into the blood and then carried to all parts of the body by blood circulation
Gas Exchange: 2 Step Process
heart (muscle) pumps blood through vessels leading to the hemocoel
step 1: exchange within environment and blood:
- gas exchange happens between the gills and the surrounding water
step 2: exchange with the blood and the individual cells
- gas exchange happens between blood in the hemocoel and the cells
- blood then returns to the gills
***in both O2 go in, and CO2 goes out